It won't be backlash or couplers because they would cause an out of round condition.
If it was a bit oversize, I would say it is tool run-out.
My 1st guess would be that comp is on but you already discredited that.
I wonder if you accidentally left "stock" in your OneCNC MOP.
I have done that before in VisualMill.
Post that section of the code so we (or you) can do the math on your interpolation.
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